This course addresses the origin of land plants and subsequent diversification of land plant vegetative and reproductive form and function. Lectures and laboratory tutorials synthesize morphological and anatomical knowledge from living organisms and fossil records with cellular, physiological and molecular information of land plants and their ancestors throughout geological time. Topics of study include the evolution of (i) vegetative and reproductive meristems, (ii) stem, leaf and root architecture, (iii) vascular tissue, (iv) the ovule habit, and (v) fertilization. (Lab materials fee: $26).